The HYDRUS software packages have become standard tools for addressing soil, agricultural, environmental, and hydrological problems involving various physical, chemical, and biological processes in the subsurface. This paper provides an overview of the capabilities of the latest Version 5 of HYDRUS, focusing on features implemented since 2016. It describes the standard and nonstandard specialized add-on modules, which significantly expand the software's capabilities. The standard add-on modules include HPx, UNSATCHEM, Wetland, Furrow, PFAS, COSMIC, DPU, and Particle Tracking. Recent developments in the HYDRUS Package for MODFLOW and additional capabilities in the graphical user interface are also discussed. The paper reviews new or improved options for simulating the fate and transport of environmental isotopes, multi-cropping systems, compensated root water uptake, and hydraulic redistribution within the rootzone. Selected applications of the HYDRUS models, such as evaluations of irrigation, low-impact development (LID), and managed aquifer recharge (MAR) schemes, are also reviewed. The paper concludes with a discussion of the use of HYDRUS in educational settings and available textbooks.The HYDRUS software packages have become standard tools for addressing soil, agricultural, environmental, and hydrological problems involving various physical, chemical, and biological processes in the subsurface. This paper provides an overview of the capabilities of the latest Version 5 of HYDRUS, focusing on features implemented since 2016. It describes the standard and nonstandard specialized add-on modules, which significantly expand the software's capabilities. The standard add-on modules include HPx, UNSATCHEM, Wetland, Furrow, PFAS, COSMIC, DPU, and Particle Tracking. Recent developments in the HYDRUS Package for MODFLOW and additional capabilities in the graphical user interface are also discussed. The paper reviews new or improved options for simulating the fate and transport of environmental isotopes, multi-cropping systems, compensated root water uptake, and hydraulic redistribution within the rootzone. Selected applications of the HYDRUS models, such as evaluations of irrigation, low-impact development (LID), and managed aquifer recharge (MAR) schemes, are also reviewed. The paper concludes with a discussion of the use of HYDRUS in educational settings and available textbooks.