Pollen nutrition plays a critical role in shaping interactions between bees and plants, influencing foraging behavior, community structure, and conservation strategies. This study investigates how variations in pollen macronutrient content (protein and lipid ratios) among 109 co-flowering plant species affect the visitation patterns of 75 subgenera of pollen-collecting bees in the Great Basin/Eastern Sierra region. The research reveals that pollen nutrition significantly predicts similarities in bee visitor communities, even after accounting for floral morphology and phylogeny. Pollen nutrition also helps delineate distinct, interconnected groups of bee subgenera and plant genera, revealing potential nutritional niches. However, variation in pollen nutrition alone does not predict bee visitor diversity, suggesting that plants offering complementary pollen nutrition may be equally valuable in supporting bee diversity. The study highlights the importance of considering pollen nutrition when selecting plants for habitat restoration, as nutritional diversity is essential for supporting diverse bee communities. The findings suggest that plants with different pollen nutritional profiles may differ in their levels of specialization or generalization within interaction networks. The research also demonstrates that pollen nutritional content and floral morphology are often correlated and both influence bee-flower associations, but nutritional content predicts bee communities when controlling for morphology. The study further shows that the most frequently interacting groups of bees and plants represent nutritional niches characterized by plants differing in average pollen P:L ratios. Additionally, pollen macronutrient content does not predict floral generalization or strength in connecting the community, indicating that plants with different pollen nutritional values are equally important in supporting diverse bee communities. The results emphasize the need for a nutritionally explicit perspective in understanding plant-pollinator interactions and informing conservation efforts. The study underscores the importance of considering pollen nutrition in habitat restoration and conservation strategies to support diverse bee communities.Pollen nutrition plays a critical role in shaping interactions between bees and plants, influencing foraging behavior, community structure, and conservation strategies. This study investigates how variations in pollen macronutrient content (protein and lipid ratios) among 109 co-flowering plant species affect the visitation patterns of 75 subgenera of pollen-collecting bees in the Great Basin/Eastern Sierra region. The research reveals that pollen nutrition significantly predicts similarities in bee visitor communities, even after accounting for floral morphology and phylogeny. Pollen nutrition also helps delineate distinct, interconnected groups of bee subgenera and plant genera, revealing potential nutritional niches. However, variation in pollen nutrition alone does not predict bee visitor diversity, suggesting that plants offering complementary pollen nutrition may be equally valuable in supporting bee diversity. The study highlights the importance of considering pollen nutrition when selecting plants for habitat restoration, as nutritional diversity is essential for supporting diverse bee communities. The findings suggest that plants with different pollen nutritional profiles may differ in their levels of specialization or generalization within interaction networks. The research also demonstrates that pollen nutritional content and floral morphology are often correlated and both influence bee-flower associations, but nutritional content predicts bee communities when controlling for morphology. The study further shows that the most frequently interacting groups of bees and plants represent nutritional niches characterized by plants differing in average pollen P:L ratios. Additionally, pollen macronutrient content does not predict floral generalization or strength in connecting the community, indicating that plants with different pollen nutritional values are equally important in supporting diverse bee communities. The results emphasize the need for a nutritionally explicit perspective in understanding plant-pollinator interactions and informing conservation efforts. The study underscores the importance of considering pollen nutrition in habitat restoration and conservation strategies to support diverse bee communities.