This section explores the question of whether generalist ancestors can give rise to specialized descendants. It delves into the mechanisms and conditions under which niche specialization can occur, highlighting the importance of environmental changes and resource availability in driving these evolutionary shifts. The discussion emphasizes the repeatability of niche spread rules, suggesting that certain patterns of adaptation are more likely to be repeated across different species and environments.This section explores the question of whether generalist ancestors can give rise to specialized descendants. It delves into the mechanisms and conditions under which niche specialization can occur, highlighting the importance of environmental changes and resource availability in driving these evolutionary shifts. The discussion emphasizes the repeatability of niche spread rules, suggesting that certain patterns of adaptation are more likely to be repeated across different species and environments.