The Myb73–GDPD2–GA2ox1 transcriptional regulatory module confers phosphate deficiency tolerance in soybean

The Myb73–GDPD2–GA2ox1 transcriptional regulatory module confers phosphate deficiency tolerance in soybean

2024 | Dandan Hu, Ruifan Cui, Ke Wang, Yuming Yang, Ruiyang Wang, Hongqing Zhu, Mengshi He, Yukun Fan, Le Wang, Li Wang, Shanshan Chu, Jinyu Zhang, Shanshan Zhang, Yifei Yang, Xuhao Zhai, Haiyan Lu, Dandan Zhang, Jinshe Wang, Fanjiang Kong, Deyue Yu, Hengyou Zhang, Dan Zhang
The study identified a major quantitative trait locus (*qPE19*) controlling seven low-phosphate (LP)-related traits in soybean (*Glycine max*) through linkage mapping and genome-wide association studies (GWAS). The gene responsible for *qPE19* was identified as *GLYCEROPHOSPHORYL DIESTER PHOSPHODIESTERASE2* (*GmGDPD2*), with haplotype 5 representing the optimal allele for LP tolerance. Overexpression of *GmGDPD2* significantly improved root architecture, phosphate efficiency, and yield-related traits, while CRISPR/Cas9-edited plants showed decreased performance. *GmMyb73* negatively regulates *GmGDPD2* by directly binding to its promoter, and *GmGDPD2* physically interacts with gibberellin 2-oxidase 1 (*GmGA2ox1*) in the plasma membrane. Overexpression of *GmGA2ox1* enhanced LP-associated traits similar to *GmGDPD2*. Double mutants for *GmGDPD2* and *GmGA2ox1* demonstrated that *GmGDPD2* regulates LP tolerance by influencing auxin and gibberellin dose-associated cell division in the root. These findings reveal a regulatory module, the Myb73–GDPD2–GA2ox1, that plays a major role in regulating LP tolerance in soybeans and has potential for developing phosphate-efficient varieties to enhance soybean production, particularly in phosphate-deficient soils.The study identified a major quantitative trait locus (*qPE19*) controlling seven low-phosphate (LP)-related traits in soybean (*Glycine max*) through linkage mapping and genome-wide association studies (GWAS). The gene responsible for *qPE19* was identified as *GLYCEROPHOSPHORYL DIESTER PHOSPHODIESTERASE2* (*GmGDPD2*), with haplotype 5 representing the optimal allele for LP tolerance. Overexpression of *GmGDPD2* significantly improved root architecture, phosphate efficiency, and yield-related traits, while CRISPR/Cas9-edited plants showed decreased performance. *GmMyb73* negatively regulates *GmGDPD2* by directly binding to its promoter, and *GmGDPD2* physically interacts with gibberellin 2-oxidase 1 (*GmGA2ox1*) in the plasma membrane. Overexpression of *GmGA2ox1* enhanced LP-associated traits similar to *GmGDPD2*. Double mutants for *GmGDPD2* and *GmGA2ox1* demonstrated that *GmGDPD2* regulates LP tolerance by influencing auxin and gibberellin dose-associated cell division in the root. These findings reveal a regulatory module, the Myb73–GDPD2–GA2ox1, that plays a major role in regulating LP tolerance in soybeans and has potential for developing phosphate-efficient varieties to enhance soybean production, particularly in phosphate-deficient soils.
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