TimeTree is a public resource that provides divergence times for species based on molecular sequence data. It allows users to access divergence times between species, timetrees of groups of species, and timelines tracing species evolution. The resource has been expanded significantly, now including over 97,000 species and over 3,000 studies. The new version integrates geological and environmental events with divergence times, enhancing understanding of the relationship between biosphere changes and species diversity. The TimeTree resource is available online at http://www.timetree.org. The TimeTree project addresses the challenge of accessing divergence time information from the rapidly growing peer-reviewed literature. It collects and synthesizes divergence time estimates from published studies, making them accessible to scientists, educators, and the public. The resource includes a multi-tiered web architecture that allows users to search for divergence times, view timetrees, and explore timelines. The new version also includes data on asteroid impacts, solar luminosity, and atmospheric composition, displayed in timepanels alongside divergence times. The TimeTree resource provides three main views: TIMETREE, TIMELINE, and NODETIME. The TIMETREE view allows users to view a timetree for a specific group of species, with timepanels showing geological and environmental events. The TIMELINE view provides a timeline of evolutionary history for a given taxon, while the NODETIME view provides divergence time estimates between two species. The resource also allows users to customize timetrees by specifying a list of species or by uploading a text file with species names. The TimeTree resource has been updated to include more detailed information, such as confidence intervals for divergence times and links to relevant studies. It also allows users to export timetrees in various formats, including Newick and image files. The resource is continuously expanding, with hundreds of new studies added each year to the Tree of Life (TTOL). The TimeTree resource is a valuable tool for researchers in evolutionary biology, molecular evolution, paleontology, and geology, providing a comprehensive database of divergence times and related information.TimeTree is a public resource that provides divergence times for species based on molecular sequence data. It allows users to access divergence times between species, timetrees of groups of species, and timelines tracing species evolution. The resource has been expanded significantly, now including over 97,000 species and over 3,000 studies. The new version integrates geological and environmental events with divergence times, enhancing understanding of the relationship between biosphere changes and species diversity. The TimeTree resource is available online at http://www.timetree.org. The TimeTree project addresses the challenge of accessing divergence time information from the rapidly growing peer-reviewed literature. It collects and synthesizes divergence time estimates from published studies, making them accessible to scientists, educators, and the public. The resource includes a multi-tiered web architecture that allows users to search for divergence times, view timetrees, and explore timelines. The new version also includes data on asteroid impacts, solar luminosity, and atmospheric composition, displayed in timepanels alongside divergence times. The TimeTree resource provides three main views: TIMETREE, TIMELINE, and NODETIME. The TIMETREE view allows users to view a timetree for a specific group of species, with timepanels showing geological and environmental events. The TIMELINE view provides a timeline of evolutionary history for a given taxon, while the NODETIME view provides divergence time estimates between two species. The resource also allows users to customize timetrees by specifying a list of species or by uploading a text file with species names. The TimeTree resource has been updated to include more detailed information, such as confidence intervals for divergence times and links to relevant studies. It also allows users to export timetrees in various formats, including Newick and image files. The resource is continuously expanding, with hundreds of new studies added each year to the Tree of Life (TTOL). The TimeTree resource is a valuable tool for researchers in evolutionary biology, molecular evolution, paleontology, and geology, providing a comprehensive database of divergence times and related information.